Nylin, S., K. Nyblom, F. Ronquist, N. Janz, J. Belicek, and M. Kallersjo. 2001. Phylogeny of Polygonia, Nymphalis and related butterflies (Lepidoptera: Nymphalidae): a total-evidence analysis. Zoological Journal of the Linnean Society 137(4): 441-468.